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Ursine colobus : ウィキペディア英語版
Ursine colobus

The ursine colobus (''Colobus vellerosus'', white-thighed colobus, Geoffroy's black-and-white colobus, white-thighed black-and-white colobus〔) is a species of primate in the Cercopithecidae family. It is found in Benin, Ivory Coast, Ghana, Nigeria, and Togo, as well as very southernmost Burkina Faso.〔Detailed range map at ("Range map: ''Colobus vellerosus''" ) 10 January 2008, International Union for Conservation of Nature (IUCN)〕 Its natural habitat is subtropical or tropical moist lowland forests. It is threatened by habitat loss.〔
==Nomenclature==
Geoffroy's black-and-white colobus was formerly thought to be a variety of ''Colobus polykomos'' with which it interbred;〔Groves, Colin P.; Angst, R. and Westwood, C. (1993) "The Status of ''Colobus polykomos'' dollmani" ''International Journal of Primatology'' 14: pp. 573–586〕 however, it has since been recognized as indeed a separate species.〔Oates, J. F. and Trocco, T. F. (1983) "Taxonomy and phylogeny of black-and-white colobus monkeys: inferences from an analysis of loud call variation" ''Folia Primatologica'' 40: pp. 83–113〕〔"Combining this information (loud calls ) with data on cranial morphology (e.g. Hull, 1979) and pelage (e.g. Rahm, 1970), Oates and Trocco, (1983) concluded that the bIack-and-white colobus monkeys are best regarded as belonging to five species: ''Colobus polykomos'' (Zimmerman, 1780); ''C. vellerosus'' (Geoffroy, 1831); ''C. guereza'' (Ruppell, 1835); ''C. satanas'' (Waterhouse, 1838); and ''C. angolensis'' (Sclater, 1860). Oates and Trocco (1983) also concluded that ''C. salanas'' retains more primitive features than other members of the group. and that ''C. guereza'' and ''C. vellerosus'', in their low-frequency loud call and other features, are the most derived species." (page 432 ) of Oates, John F.; Bocian, C. M. and Terranova, C. J. (2000) "The loud calls of black-and-white colobus monkeys: their adaptive and taxonomic significance in light of new data" , ''In'' Whitehead, Paul F. Jolly, Clifford J. (2000) ''Old World Monkeys'' Cambridge University Press, Cambridge, England, pp. 431–452, ISBN 978-0-521-02809-7〕

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